Typhlotriton is an incipient blind salamander living in the caves of southwestern Missouri. It detects its food by the sense of touch without the use of its eyes. It is stereotropic. Its eyes show the early stages in the steps of degeneration from those of salamanders living in the open to those of the degenerate Typhlomolge from the caves of Texas. The lids are in process of obliteration, the upper overlapping the lower so that the eye is always covered in the adult. The sclera possesses a cartilaginous band in the larval stages but not in the adult. The disappearance of the cartilage is probably an incident of the metamorphosis, not of the degeneration the eye is undergoing. The lens is normal. The retina is normal in the larva with a proportionally thicker ganglionic layer than in the related epigæan forms. Marked ontogenic degenerations take place during and shortly after the metamorphosis. a. The outer reticular layer disappears. b. The rods and cones lose their complexity of structure, such as differentiation into inner and outer segments, and finally are lost altogether.