Four New Genera of Leptophlebiidae (Ephemeroptera: Atalophlebiinae) from Southern South America

Abstract
Four new monotypic genera of Leptophlebiidae, Archethraulodes, Demoulinellus, Rhigotopus and Secochela are established. Four new species are described. Three, Archethraulodes spatulus, Rhigotopus andinensis and Secochela illiesi are described from imagines. One, Demoulinellus coloratus is known from both imagines and nymph. Descriptions of the eggs of genera are included.