In recent publications we have described the distribution and structure of vascular transfer cells in the floral axes of wheat (Zee and O'Brien 1971) and the presence of a xylem discontinuity composed of modified tracheary elements at the base of the peri carp (Zee and O'Brien 1970a). We have also examined the structure of the pigment strand during grain formation (Zee and O'Brien 1970b). In order to find out if these structures are present in the florets of a non-festucoid grass we have examined a member of the Paniceae, Japanese millet (Echinochloa utilis Ohwi & Yabuno).