Abstract
The effect of 2 levels of thinning (zero and 2/3 of basal area removed) and 3 levels of N fertilization (0, 224 and 448 kg N/ha) on crown development of codominant, 24-yr old Douglas-fir (P. menziesii (Mirb.) Franco) trees was studied over a 5-7 yr period. Thinning and heavy fertilization separately increased needle mass per tree after 7 yr by 90% and, when combined, by 271%. Yearly needle production peaked 2-3 yr after fertilization and resulted from an increase in needle size, needle number of shoots produced. Maximum foliage mass per tree was reached 4-7 yr after fertilization. Thinning effect on needle production was lower initially, but increased throughout the study period. A continuing foliage production in branches low in the crown contributed to thinning effect on foliage mass and crown size. Foliage distribution was affected most in the top half of the crown by fertilization and in the bottom half of the crown by thinning. Fertilization increased branch elongation at all crown heights, but thinning alone had no effect on crown width down to whorl 12.

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