Abstract
Lodging results from low dry matter per unit length of culm; this may result from a relatively low composition of lignin and various reserve materials, such as disaccharides and polysaccharides, or from the development of relatively small slender culms. Culms, small in diam. but medium or high in composition of carbohydrates, may result from direct causes such as thick seedlings or the planting of small seed. Such culms, however, exhibit a low dry matter content per unit length of culm. Culms low in composition of carbohydrates are due to the development within the plant of a low carbohydrate N ratio, which may result from hypernutrition, shading, or relatively high temp. Under field conditions lodging results from the interaction of all these environmental factors. Hypernutrition not only tends to induce greater elongation of culms, but through a stimulation of the tillering processes, it increases the thickness of stand and hence results in a greater degree of shading around the base of the plants. In warm, rainy or cloudy seasons, therefore, the conditions are most favorable for the development of lodging on rich soils.

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