Abstract
The Lower Carboniferous seed Samaropsis scotica Calder has been re-investigated from new material and assigned to a new genus Lyrasperma. It is shown to have a nucellar apex with a pollen chamber and a funnel-like structure interpreted as an expanded salpinx. The free parts of the two integumental lobes diverge as apical horns; there is therefore no micropyle. Eurystoma angulare gen. et sp. nov. possesses a similar nucellar apex but is usually quadrangular in section and has four apical lobes. The new genera are placed in the group Lagenostomales although the expanded salpinx is a character new to that group.