WOODPECKER PREDATION ON BARK BEETLES IN ENGELMANN SPRUCE LOGS AS RELATED TO STAND DENSITY

Abstract
Eighty Engelmann spruce logs (Picea engelmannii Parry), which were distributed in open (bulldozed to clearcut), semi-open (selectively cut), and dense (uncut) spruce forest, were infested with Dendroctonus obesus (Mann.) and Ips pilifrons Sw. and fed upon by northern three-toed and hairy woodpeckers. Sections of the infested boles were covered with screen to prevent woodpecker feeding and all logs were left in the field over one winter and through the following summer. By spring, the spruce beetle brood was reduced approximately 50% in all three forest areas. By fall, the spruce beetle brood suffered a 71, 83 and 52% reduction in the open, semi-open and dense forest, respectively. Spruce beetles appeared to be in greatest numbers in the semi-open forest, resulting in the greatest woodpecker predation there. Woodpeckers did not feed in the open meadows. Estimates in the spring, before the Ips emerged from the logs, showed that woodpeckers reduced the Ips brood by 76 and 11% in the open and semi-open forest, respectively. Ips were not found in dense forest or in meadows.