Seasonal History of the Balsam Woolly Aphid in the Pacific Northwest

Abstract
The balsam woolly aphid, Chermes (Adelges) piceae Ratz., is widely distributed in Europe and North America. It infests many species of Abies (true firs) and may be found attacking its hosts at elevations from near sea level to timberline (5,000 to 6,000 feet). Because of the great range of environments inhabited by the balsam woolly aphid, its biology differs considerably from one region to another. Perhaps the most variable feature in its biology is seasonal history. Karafiat and Franz (1956) reported three generations per year in central Europe, whereas Varty (1956) noted two and sometimes a partial third generation in Scotland. Balch (1952) found only two generations in the Maritime Provinces of Canada, but Amman noted three and sometimes a partial fourth generation in southeastern United States.