Introduction of the D-genome chromosomes from bread wheat into hexaploid triticale with a complete rye genome

Abstract
Hexaploid triticale (× Triticosecale Wittmack) lines selected from the progeny of octoploid × tetraploid triticale hybrids were karyotyped using C-banding. The number of D-genome chromosome pairs substituted for A- and (or) B-genome chromosomes ranged from 0 to 4, averaging 2.1 substitutions per line. Every D-genome chromosome was present in at least 1 of the 70 lines analyzed. The most frequent were chromosomes 3D and 6D, followed by 1D. Of the 14 possible substitutions, 12 were present in the homozygous condition, 1 (4D/4B) was still segregating, and 6D/6B was absent. With the exception of one 1D/1R substitution and one 7RS/4DS translocation, all lines had a complete rye genome. Key words: triticale, chromosome substitutions, D genome.